Allosaurus head
Hello everyone. Here is a drawing of the head of Allosaurus fragilis, the top predator of the Morrison Formation of Late Jurassic North America. This drawing has been on my to-do list for quite some time, and I’m happy that it’s finally finished. The drawing was made with No. 2 pencil on printer paper.
May 9, 11, and 13 – Rest in Peace: The Lemuria Festival of the Dead
Do you believe in ghosts? The ancient Romans certainly did. The spirits of the Undead were a real concern and a real fear for the ancient Romans. Therefore, it was important that these otherworldly beings be kept happy and pacified as much as possible.
Many people nowadays associate all things spooky with October 31, Halloween. You might be interested to know that the ancient Romans, too, had their own version of Halloween, except it occurred in May instead of October and it lasted for three days instead of just one. It was known as the Lemuria, named after the lemures, the restless malevolent spirits of the dead. These formless shapeless wraiths might be haunting you for a variety of reasons: they were not given a proper burial, they want revenge for a wrong committed upon them, or any number of things. Rituals were conducted to drive ghosts out of your home, and offerings were left outside homes so that the ghosts could be appeased and leave the family alone. Sounds similar to trick-or-treating, doesn’t it?
The Lemuria was held on May 9, 11, and 13 – notice that it skipped over May 10 and May 12. Because of the Lemuria festival, it was believed that May was an unlucky month to get married; any couple in love who wished to marry would have to wait until June. All temples were also closed on these three days (Ovid, Fasti, book 5, May 9).
The day was originally held in honor of Remus, Romulus’ twin brother who was murdered when a dispute arose between the two over who should be the first king of the settlement which they had established on the bank of the Tiber. May 9 was originally known as the Remuria, the remembrance feast of Remus and for all other fallen spirits of a person’s family. Over time, the first R in Remuria changed to an L. Eventually “the silent spirits”, as they were known, were collectively referred to as lemures. They were, in essence, ancient Roman poltergeists (Ovid, Fasti, book 5, May 9). The lemures are generally distinguished from the manes as being more hostile and also more likely to haunt people’s homes. Perhaps this is the reason why the word “lemures” was a synonym for “larva” (William Warde Fowler, The Roman Festivals of the Period of the Republic: An Introduction to the Study of the Religion of the Romans. London: Macmillan and Co., Ltd., 1899. Page 108), because the larvae of insects make homes for themselves inside the bodies of plants or other animals. A beetle’s larva might burrow into the bark of a tree, or a parasitic wasp’s larva might develop inside an unsuspecting host. These creatures nest themselves within other homes, as the lemures might unexpectedly make a new home for itself inside your own home, or possibly within you personally.
In his 1899 overview of ancient Roman religious festivals, William W. Fowler posits that the Lemuria, along with the earlier Feralia festival conducted in late February, might be one of the most archaic of Roman rituals, conducted at a time when primitive cultures feared demons and undead spirits and needed to periodically expel them (William Warde Fowler, The Roman Festivals of the Period of the Republic: An Introduction to the Study of the Religion of the Romans. London: Macmillan and Co., Ltd., 1899. Page 107).
Fowler also proposes that the Lemuria would have hit many Romans much closer to home than the Feralia would have. The dead transformed into lemures under many circumstances – violent death, suicide, bodies not buried properly or not buried at all, wrongs that were not avenged, and other reasons – and all of these were frequent if not daily occurrences in the ancient world. Maybe that’s part of the reason why you needed three whole days to placate any irate entities and protect your family from harm, because there were A LOT of angry bitter pissed-off ghosts out there (William Warde Fowler, The Roman Festivals of the Period of the Republic: An Introduction to the Study of the Religion of the Romans. London: Macmillan and Co., Ltd., 1899. Pages 107-109).
Alright, enough of the background information. Time now to get into the details of how this festival was carried out. The poet Ovid provides us with the majority of information regarding the rituals of this spooky time of year. On May 9, the head of the household (always a man) would rise from his bed and began the necessary rites needed to placate any hostile spirits that may wish him, his family, or his property harm. He would go outside barefoot and walk around his house nine times, all the while tossing black beans over his shoulder. Black was the color that was associated with Underworld entities, and it was believed that they were attracted towards food that was black in color. While the man of the house was tossing the black beans outside, he would repeat the incantation “I throw these. With these beans, I redeem me and mine”. When the man of the house had performed this ritual nine times, he again washed his hands and rang a bronze bell saying “Ancestral spirits, depart!” – with this act, the sacred rites are concluded (Ovid, Fasti, book 5, May 9).
The part about ringing the bell and calling the ghosts who were haunting his home to leave immediately sounds similar to the “wassail” ritual of making noise to drive evil spirits away from apple orchards. It’s also similar to ideas held by some tribes that demons and evil spirits are driven away by excessive noise (A Merry Tudor Christmas; Lucy Worsley’s Christmas Carol Odyssey; Victorian Farm Christmas, episode 3; Edwardian Farm, episode 5).
Hopefully, all of these methods would achieve the desired result. However, if you were an ancient Roman, and you suspected that an evil spirit had entered your house, and you performed the proscribed exorcism rituals, and you still heard things go “bump” in the night…then you were in big trouble.
Sources:
- Ovid, Fasti, book 5, May 9. https://www.poetryintranslation.com/PITBR/Latin/OvidFastiBkFive.php.
- Fowler, William Warde. The Roman Festivals of the Period of the Republic: An Introduction to the Study of the Religion of the Romans. London: Macmillan and Co., Ltd., 1899.
- A Merry Tudor Christmas. Hosted by Lucy Worsley. BBC, 2019.
- Edwardian Farm. Episode 5. BBC, 2010.
- Lucy Worsley’s Christmas Carol Odyssey. Hosted by Lucy Worsley. BBC, 2019.
- Victorian Farm Christmas. Episode 3. BBC, 2009.
May 15 – The Feast of Mercury
May 15 was the date of the Mercuralia, the Feast of Mercury. Mercury was the Roman version for the ancient Greek god Hermes, the messenger of the gods and a bringer of dreams, and the patron god of messengers, tourists, travelling merchants, as well as of thieves and game-cheaters (H. A. Guerber, Myths of Greece and Rome. New York: American Book Company, 1893. Page 134).
The ancient Greek god Hermes, and his Roman counterpart Mercury, have curious origins and legends attached to them. Although they are often given the designation of being a divine messenger, both Hermes and Mercury seem to have started off as rain gods who also had some connection to the Underworld. According to Samuel F, Dunlap, a 19th Century “theologian” (I use that term EXTREMELY loosely, since his writings bear more of a resemblance to the rambling rantings of a religious crack-pot or cult leader), the name Hermes comes from Haram-eias, who might he related to Baal-Ram. The name Mercury is related to the Phoenician rain god Mar, and also held the title Mar-Kuri, “Mar of the Dead”. In Greek myth, Hermes was one of several sky gods which included Zeus, Apollo, and Helios. Hermes is mentioned as a rain god who nourished the earth with water from Heaven, and who possibly gave restorative power to the dead. The rooster was an animal sacred to Hermes and served as his symbol (Samuel Fales Dunlap, Sōd: The Mysteries of Adoni. London: Williams and Norgate, 1861. Pages 77-78).
The reason why the Feast of Mercury takes place on May 15 is because his first temple in the city of Rome was dedicated on this day, according to the Roman historian Livy. According to his report, the first temple of Mercury was officially opened on May 15, 495 BC (Gary Forsythe, The historian L. Calpurnius Piso Frugi and the Roman Annalistic Tradition. Berkeley: The University of California Press, 1994. Page 147).
However, it’s possible that May 15 wasn’t just some random date; the date of the opening for his temple may have been deliberately timed to take place on this day. There are two reasons why I say this. Firstly, the “ides” were regarded by the ancient Romans has being very important dates within the calendar. According to Roman myth, Mercury was the son of the goddess Maia, of whom the month of May is named after (H. A. Guerber, Myths of Greece and Rome. New York: American Book Company, 1893. Page 131). Therefore, having the temple’s dedication take place on the Ides of May would have been appropriate, considering Mercury’s divine parentage.
Secondly, it’s possible that Feast of Mercury takes place on May 15 as a continuation of the Lemuria festival. The Lemuria was a festival dedicated to the dead (one of several in the ancient Roman calendar) which was celebrated on May 9, 11, and 13. According to Roman mythology, the god Mercury had a part to play in things related to the souls of the departed: “To Mercury was intrusted (sic) the charge of conducting the souls of the departed to Hades” (H. A. Guerber, Myths of Greece and Rome. New York: American Book Company, 1893. Page 137).
But by and large, Mercury was not thought of as a god of death. Rather, he was associated with commerce, news, dreams, and cleverness. “The profession of merchandise (saith Plutarch) was honourable, as it brought home the produce of barbarous countries, engaged the friendship of kings, and opened a wide field of knowledge and experience” (Anonymous, The Anniversary Calendar, Natal Book, and Universal Mirror. Volume II. London: William Kidd, 1832. Page 485).
On May 15, Roman merchants would take water from the well of Porta Capena, a well that was believed to be sacred to Mercury, and sprinkle it on themselves, their ships, and their cargo to protect them while travelling (C. Scott Littleton, ed., Gods, Goddesses, and Mythology. Volume 6: Inca-Mercury. Tarrytown: Marshall Cavendish, 2005. Page 861). The water in Mercury’s well was known as acqua Mercurii was believed to aid in forgiving sins – both those committed in the past as well as any that might be committed in the future – and was thought to bring good luck. Travelling merchants needed all the luck that they could when carrying out commerce. The threats of storms, shipwrecks, pirates, thieves, bandit gangs, and even outbreaks of war were ever-present on their minds (Rebecca I. Denova, Greek and Roman Religions. Hoboken: John Wiley & Sons, Inc., 2019. Page 131).
The Romans also had a second Mercuralia festival later in the year, and this one lasted for more than just one day. The second Mercuralia was a six-day-long celebration that lasted from July 14 to 19 (Anonymous, The Anniversary Calendar, Natal Book, and Universal Mirror. Volume II. London: William Kidd, 1832. Page 485). We know a little bit more about the celebrations that took place during this period than the earlier festival in the middle of May. A sow was sacrificed, and, according to the Greek writer Athenaeus, “They poured libations at the conclusion of dinner and offered them to Hermes, not, as in later times, to Zeus the Fulfiller. For Hermes is regarded as the patron of sleep. So they pour the libation to him also when the tongues of the animals are cut out on leaving a dinner. Tongues are sacred to him because he is the god of eloquence” (Athenaeus, Deipnosophistae, excerpts from Book 1, 16B-C; Samuel Fales Dunlap, Sōd: The Mysteries of Adoni. London: Williams and Norgate, 1861. Pages 77).
The symbol of both Hermes and Mercury was the caduceus. Myth says it was presented to Mercury as a gift from Apollo as a reward for inventing the lyre (H. A. Guerber, Myths of Greece and Rome. New York: American Book Company, 1893. Page 134). As an interesting coincidence (or perhaps it isn’t a coincidence), the third full week in May is unofficially known as “National Emergency Medical Service Week” (“National EMS Week”).
Sources:
- Anonymous. The Anniversary Calendar, Natal Book, and Universal Mirror. Volume II. London: William Kidd, 1832.
- Athenaeus. Deipnosophistae, excerpts from Book 1, 16B-C. http://penelope.uchicago.edu/Thayer/E/Roman/Texts/Athenaeus/home.html.
- Denova, Rebecca I. Greek and Roman Religions. Hoboken: John Wiley & Sons, Inc., 2019.
- Dunlap. Samuel Fales. Sōd: The Mysteries of Adoni. London: Williams and Norgate, 1861.
- Forsythe, Gary. The historian L. Calpurnius Piso Frugi and the Roman Annalistic Tradition. Berkeley: The University of California Press, 1994.
- Guerber, H. A. Myths of Greece and Rome. New York: American Book Company, 1893.
- Littleton, C. Scott, ed. Gods, Goddesses, and Mythology. Volume 6: Inca-Mercury. Tarrytown: Marshall Cavendish, 2005.
- National Day. “National EMS Week”. https://nationaldaycalendar.com/national-ems-week-third-full-week-of-may/.
Evidence of Therizinosaurs in North America during the Late Cretaceous Period
Introduction
For many years, paleontologists have known about the presence of therizinosaurs (formerly classified as segnosaurs) in Asia, especially within what’s now Mongolia and China. However, Asia and North America were linked during a considerable portion of the Cretaceous Period, and this resulted in an interchange of faunas between the two continents, notably ceratopsians, pachycephalosaurs, tyrannosaurs, and maniraptorans. Could therizinosaurs, which had hitherto been exclusively Asian, have lived in North America as well?
A pair of Tarbosaurus attacking a herd of Therizinosaurus somewhere in Mongolia, approximately 80 million years ago. © Gregory S. Paul (1988). Image used with permission.
During the early 2000s, that question was answered with a definitive “yes”. Two genera of therizinosaurs have been described from North America, named Falcarius and Nothronychus. Falcarius represents possibly the earliest stage in therizinosaur evolution, dated to the early Cretaceous Period, while Nothronychus is much larger and more advanced and is dated to the middle Cretaceous. The presence of these two creatures clearly shows that therizinosaurs existed in North America, but so far they have only been found in rocks dated to the early and middle parts of the Cretaceous Period. One wonders if therizinosaurs managed to stay in North America right up until the end of the Mesozoic, 66 million years ago. Would they have kept evolving, becoming larger and more advanced? Would they have lived alongside Triceratops and Tyrannosaurus? (1)
It just so happens that there are a few pieces of evidence here and there which suggest that therizinosaurs did survive past the middle Cretaceous within North America, and that they kept living in North America up to the end of the Cretaceous Period.
The Evidence
The idea that there were therizinosaurs in late Cretaceous North America was first proposed by the German paleontologist Hans Sues in 1978. Specifically, he was writing about a particular specimen that had been uncovered in the Dinosaur Park Formation, located in Alberta, Canada, in rocks dated to the Campanian Stage of the Cretaceous Period. The specimen in question was a single “frontal” bone, which forms part of the skull. Today, this specimen is in the collections of the Carnegie Museum of Natural History, categorized as “CMN 12355” (NOT 12349 as you’ll sometimes see in internet searches). In his paper, Sues thought that this frontal bone belonged to a “raptor” dinosaur, and listed it as “gen. et sp. indet.”, which is an abbreviated Latin way of saying “genus and species undetermined” (2).
“CMN 12355”: A frontal bone which may belong to a therizinosaur. Left top: ventral view. Right top: dorsal view. Left bottom: lateral view. Right bottom: medial view. © Tracy Ford. Image from Paleofile.com. Used with permission. http://www.paleofile.com/Dinosaurs/Theropods/Segnosaurincertae.asp
Saying that this bone belonged to a raptor is understandable, since the dromaeosaurs and the therizinosaurs are related to each other. Both groups are located in a clade called the “maniraptorans”, which includes the ornithomimids, the oviraptorosaurs, the therizinosaurs, and famously, the dromaeosaurs and troodontids – the so-called “raptors” with their famous killing claws.
The second piece of evidence came in the early to mid 1980s. A single bone called an “astragalus”, which forms part of the ankle, was found in the Hell Creek Formation in rocks dated to the very end of the Cretaceous Period. In 1984, the Canadian paleontologist Dale Russell listed this single peculiar find in a long list of specimens uncovered in the Hell Creek Formation during the middle 1980s. However, this particular specimen has never been analyzed or described in a publication exclusively devoted to this bone. It is simply listed as “therizinosaurid indet.”. In 1992, Kenneth Carpenter looked at this bone, and concluded that it actually belonged to Tyrannosaurus, not a therizinosaur (3).
In 1987, the Canadian paleontologist Philip Currie, who is widely acknowledged as the world’s expert on meat-eating dinosaurs, took a second look at the frontal bone which Sues had examined in the late 1970s, and concluded that Hans Sues had made a mistake. It wasn’t a raptor, but was instead a “segnosaur”, which was the way therizinosaurs were called back then. Currie stated that the bone looked similar to the frontal bone of an Asian therizinosaur called Erlikosaurus, and so he reclassified the bone as “cf. Erlikosaurus” (4).
In 1992, Philip Currie did a more thorough examination of possible therizinosaur finds in Canada. He again wrote about the frontal bone which was initially described in 1978, but he also added two more specimens to the discussion table, both of which were housed in the collection of the Royal Tyrell Museum of Paleontology (RTMP). These specimens were given the identification codes “RTMP 81.16.231” (again, Currie classified this specimen as “cf. Erlikosaurus”) and “RTMP 79.15.1” (a “pedal ungual”, or foot claw, which was classified as “cf. therizinosaurid”) (5).
“RTMP 79.15.1”: A foot claw which may belong to a therizinosaur. © Tracy Ford. Image from Paleofile.com. Used with permission. http://www.paleofile.com/Dinosaurs/Theropods/Segnosaurincertae.asp
In 2001, Michael Ryan and Anthony Russell conducted their own analysis of North American therizinosaur finds. They confirmed Currie’s claim that the frontal bone found in 1978 did indeed come from a therizinosaur. They also wrote about a neck vertebra found in the Scollard Formation (specimen identification code is “RTMP 86.207.17”), which dates to the very end of the Cretaceous Period, and which they classified as “Therizinosauridae indet.” (6).
Body fossils of therizinosaurs may be rare in North America, but footprints which may belong to therizinosaurs are more abundant. The first footprints were discovered in the 1990s in the Harebell Formation of northwestern Wyoming. According to an article published in 1996, these footprints were unique because they looked like theropod prints except that they had four toes instead of three – unique among theropods, therizinosaurs have four main toes. The authors postulated that the footprints belonged to an animal whose physical remains had not yet been discovered (7).
In 2011, a single therizinosaur footprint was discovered in Denali National Park, Alaska. The rock that the footprint was found in was part of the Cantwell Formation, which spans 80-65 MYA, and the footprint was placed in a layer dated to about 71-69 MYA. Depending upon which source that you read concerning geological dating, this date of 71-69 MYA either marks the boundary between the where the Campanian Stage ends and the Maastrichtian Stage begins, or else it is the earliest phase of the Maastrichtian Stage. In 2012, Anthony R. Fiorillo of the Perot Museum of Nature and Science (located in Dallas, Texas) published an article concerning this peculiar footprint (8). You can see a photo of it here.
In 2013 and 2014, Anthony R. Fiorillo and a team of other researchers returned to the site in Denali National Park and found a total of thirty-one therizinosaur footprints, along with numerous hadrosaur footprints as well. Like the first footprint that had been found in 2011, all of the other footprints were in rock dated to 71-69 MYA. The fact that footprint trackways of both hadrosaurs and therizinosaurs were found together might indicate that these animals traveled together, possibly for mutual protection. An article was published in August 2018 detailing these discoveries (9).
Species Identification
As we have seen in the previous section, there is some evidence in the way of footprints and a handful of isolated bones which suggests that therizinosaurs inhabited North America during the late Campanian or early Maastrichtian Stages of the Cretaceous Period. However, is there any way that we can identify which particular genus or species that these fossils belong to?
The subject of identification has been especially contentious concerning the footprints that were found in Wyoming and Alaska. So far, footprints form the majority of finds that are attributed to late Cretaceous therizinosaurs within North America. The problem is that it is difficult to identify a particular genus or species based solely on footprints, unless the shape of the footprint is extremely distinctive. Another problem is that while footprints are abundant, very few body fossils have been found, and none of them are highly diagnostic. Most researchers who examined them determined vaguely that the creature was a therizinosaur, but they couldn’t be more specific than that, with the exception of Philip Currie who proposed that they might belong to Erlikosaurus or a creature very similar to it.
Because it is so difficult to match a footprint with a particular animal, paleontologists often ascribe footprints their own genus and species names. This is what is referred to as an “ichnogenus”, which is a genus of animal known only from trace fossils, such as footprints, rather than actual physical body fossils.
In the 1996 article which discussed the unusual footprints found in Wyoming, the footprints were ascribed to the ichnogenus Exallopus (pronounced as Ex-ALLO-pus, meaning “from different foot” due to its unusual shape) and its species name was given as Exallopus lovei. The type specimen is identified as “DMNH 5989”, and it was identified as a coelurosaur. According to the website Fossilworks, “Its type locality is Whetstone Creek tracksite, which is in a Maastrichtian terrestrial sandstone in the Harebell Formation of Wyoming” (10). The following year in 1997, the genus name was changed from Exallopus to Saurexallopus (SORE-ex-ALLO-pus), because the name Exallopus was already taken by a species of marine worm (11). Another species, Saurexallopus zerbsti, was named in a 2003 article. The type specimen is identified as “CUMWC 224.2”. According to Fossilworks, “Its type locality is Zerbst Ranch Tracksite, which is in a Lancian fluvial sandstone/sandstone in the Lance Formation of Wyoming” (12). In 2014, a third species was named called Saurexallopus cordata based upon a single footprint fount in British Columbia, Canada, and dated to the Wapiti Formation of the late Cretaceous Period (13).
While all of the scientific articles concerning Saurexallopus identify it as a theropod, there has been some dispute as to what particular type of theropod it is. The original article which was written in 1996 identified it as a coelurosaur. In 2012, Anthony Fiorillo and Thomas Adams identified Saurexallopus as a therizinosaur (14). In an article written in 2015, Saurexallopus was identified as an oviraptorid (15). In an article written in 2018, Saurexallopus was simply identified as a theropod without any specific affinity (16). The website Fossilworks identifies Saurexallopus as a therizinosaur (17).
Reconstructing Saurexallopus
During the late 1980s and early 1990s, Philip Currie made comparisons between the various finds in North America with the Asian species Erlikosaurus. According to a phylogenic analysis of therizinosaur genera which was conducted in 2019, Erlikosaurus was closely related to Nothronychus, a therizinosaur which lived in North America during the middle Cretaceous Period. Since Saurexallopus is believed to be physically similar to Erlikosaurus, it is likely that it was genetically related as well, and as such would have been genetically related to Nothronychus. It is therefore quite possible that Erlikosaurus, Nothronychus, and Saurexallopus would have been similar in appearance (18).
Upper jaw and right foot of the Asian therizinosaur Erlikosaurus. Saurexallopus was probably similar in appearance to this genus. Illustration from Rinchen Barsbold and Altangerel Perle (1980) “Segnosauria, a new infraorder of carnivorous dinosaurs”. Acta Palaeontologica Polonica, 25 (2): pages 187-195. https://www.app.pan.pl/article/item/app25-187.html. Creative Commons Attribution License.
We can guess that Saurexallopus reached a similar length to Erlikosaurus, measuring about fifteen to twenty feet long (Holtz claims that Erlikosaurus was smaller than other authors do, although his estimate of Nothronychus is in fitting with the size bracket mentioned above) (19). Unlike the eponymous Therizinosaurus, which possessed long scythe-like finger claws (hence its name, which translates to “scythe lizard”), Nothronychus possessed shorter hook-shaped claws, which looked very similar to the stereotypical talons that are seen on carnivorous dinosaurs like Allosaurus and Torvosaurus. These claws were only one-third the size of the claws of Therizinosaurus, but they were well-suited for pulling down branches, for digging (if they could pronate their hands, but that’s a whole other argument), and for smacking the daylights out of any would-be predator. Thomas R. Holtz Jr. has compared therizinosaurs to the large ground sloths of the Cenozoic Era, and the analogy has some merit (20). Saurexallopus and other therizinosaurs likely lived a similar lifestyle and occupied a similar ecological niche, with the possible exception of Falcarius, which may have had a more cursorial lifestyle similar to early coelurosaurs like Ornitholestes.
Based upon their place within the dinosaur family tree, as well as from fossil finds, we are fairly certain that therizinosaurs were feathered. Therefore, it is almost certain that Saurexallopus would have had some form of feather covering as well, although whether it was over the entire body or only partially cannot be determined.
Below is a drawing that I made of Saurexallopus, based upon Erlikosaurus and Nothronychus. The erect mane running down the middle of its neck, back, and tail are just artistic conjecture.
Saurexallopus. © Jason R. Abdale. May 7, 2020.
Conclusions
So where does all of this information lead us? So far, there is some evidence which suggests that therizinosaurs were living in Alberta, Canada and Alaska, USA during the late Campanian Stage or early Maastrichtian Stage of the late Cretaceous Period up until about 70 MYA or thereabouts. As such, they would have lived side-by-side with creatures such as Albertosaurus, Edmontosaurus, and Hypacrosaurus. There is only one piece of evidence, a single neck vertebra, which suggests that therizinosaurs existed in North America during the Maastrichtian Stage of the Late Cretaceous. However, no specimens that can be definitely and unquestionably identified as belonging to a therizinosaur have been found in the Hell Creek Formation. Therefore, as far as our current evidence goes, it is unlikely that therizinosaurs lived side-by-side with Triceratops and Tyrannosaurus. However, this may change in the future if more body fossils are discovered.
Sources
- Utah’s Dino Graveyard; When Dinosaurs Roamed America.
- Lindsay Elizabeth Zanno. A Taxonomic and Phylogenetic Reevaluation of Therizinosauria (Dinosauria: Theropoda): Implications for the Evolution of Maniraptora. PhD dissertation, submitted to the University of Utah. December 2008. Page 172.
- Lindsay Elizabeth Zanno. A Taxonomic and Phylogenetic Reevaluation of Therizinosauria (Dinosauria: Theropoda): Implications for the Evolution of Maniraptora. PhD dissertation, submitted to the University of Utah. December 2008. Page 172; Dinosaur Mailing List. “Re: Yet even more questions (and I’m sure there’ll be more…)”, by Mickey Mortimer (June 22, 2002). http://dml.cmnh.org/2002Jun/msg00369.html; Theropod Database. “Therizinosauroidea”. http://theropoddatabase.com/Therizinosauroidea.htm.
- Lindsay Elizabeth Zanno. A Taxonomic and Phylogenetic Reevaluation of Therizinosauria (Dinosauria: Theropoda): Implications for the Evolution of Maniraptora. PhD dissertation, submitted to the University of Utah. December 2008. Page 172.
- Lindsay Elizabeth Zanno. A Taxonomic and Phylogenetic Reevaluation of Therizinosauria (Dinosauria: Theropoda): Implications for the Evolution of Maniraptora. PhD dissertation, submitted to the University of Utah. December 2008. Page 172; Dinosaur Mailing List. “Re: Yet even more questions (and I’m sure there’ll be more…)”, by Mickey Mortimer (June 22, 2002). http://dml.cmnh.org/2002Jun/msg00369.html.
- Lindsay Elizabeth Zanno. A Taxonomic and Phylogenetic Reevaluation of Therizinosauria (Dinosauria: Theropoda): Implications for the Evolution of Maniraptora. PhD dissertation, submitted to the University of Utah. December 2008. Page 172.
- J. D. Harris, K. R. Johnson, J. Hicks and L. Tauxe (1996). “Four-toed theropod footprints and a paleomagnetic age from the Whetstone Falls Member of the Harebell Formation (Upper Cretaceous: Maastrichtian), northwestern Wyoming”. Cretaceous Research, 17: 381-401.
- Anthony R. Fiorello and Thomas L. Adams (2012). “A therizinosaur track from the Lower Cantwell Formation (Upper Cretaceous) of Denali National Park, Alaska”. Palaios, 27: 395-400.
- Anthony R. Fiorello and Thomas L. Adams (2012). “A therizinosaur track from the Lower Cantwell Formation (Upper Cretaceous) of Denali National Park, Alaska”. Palaios, 27: 395-400; “The Lower Cantwell Formation and Its Fossils”; “Therizinosaur: prehistoric predator set standard for ‘weird’ in Alaska”; “First North American co-occurrence of Hadrosaur and Therizinosaur tracks found in Alaska”.
- Fossilworks. “Saurexallopus lovei”. http://fossilworks.org/bridge.pl?a=taxonInfo&taxon_no=65844.
- J. D. Harris, K. R. Johnson, J. Hicks and L. Tauxe (1996). “Four-toed theropod footprints and a paleomagnetic age from the Whetstone Falls Member of the Harebell Formation (Upper Cretaceous: Maastrichtian), northwestern Wyoming”. Cretaceous Research, 17: 381-401; J. D. Harris (1997). “Four-toed theropod footprints and a paleomagnetic age from the Whetstone Falls Member of the Harebell Formation (Upper Cretaceous: Maastrichtian), northwestern Wyoming: a correction”. Cretaceous Research, 18: 139.
- Martin G. Lockley, G. Nadon, and Philip J. Currie. (2003). “A diverse dinosaur-bird footprint assemblage from the Lance Formation, Upper Cretaceous, eastern Wyoming; implications for ichnotaxonomy”. Ichnos, 11: 229-249; Fossilworks. “Saurexallopus zerbsti”. http://fossilworks.org/bridge.pl?a=taxonInfo&taxon_no=81011.
- R. T. McCrea, L. G. Buckley, A. G. Plint, Philip J. Currie, J. W. Haggart, C. W. Helm, and S. G. Pemberton (2014). “A review of vertebrate track-bearing formations from the Mesozoic and earliest Cenozoic of western Canada with a description of a new theropod ichnospecies and reassignment of an avian ichnogenus”. In Lockley Martin G.; Lucas, Spencer G., eds. New Mexico Museum of Natural History & Science. Bulletin 62: Fossil Footprints of Western North America. Albuquerque: New Mexico Museum of Natural History & Science, 2014. Page 87.
- Anthony R. Fiorello and Thomas L. Adams (2012). “A therizinosaur track from the Lower Cantwell Formation (Upper Cretaceous) of Denali National Park, Alaska”. Palaios, 27: 395-400.
- R. T. McCrea, D. H. Tanke, L. G. Buckley, M. G. Lockley, J. O. Farlow, L. Xing, N. A. Matthews, C. W. Helm, S. G. Pemberton and B. H. Breithaupt (2015). “Vertebrate ichnopathology: pathologies inferred from dinosaur tracks and trackways from the Mesozoic”. Ichnos, 22 (3–4): 235-260.
- Martin Lockley, Gerard Gierlinski, Lidia Adach, Bruce Schumacher, and Ken Cart (2018). “Newly Discovered Tetrapod Ichnotaxa from the Upper Cretaceous Blackhawk Formation, Utah”. In Spencer G. Lucas and Robert M. Sullivan, eds. New Mexico Museum of Natural History and Science. Fossil Record 6, Volume 2: Bulletin 79. Albuquerque: New Mexico Museum of Natural History and Science, 2018. Pages 469-480.
- Fossilworks. “Saurexallopus”. http://fossilworks.org/bridge.pl?a=taxonInfo&taxon_no=65843.
- Scott Hartman, Mickey Mortimer, William R. Wahl, Dean R. Lomax, Jessica Lippincott, and David M. Lovelace (2019). “A new paravian dinosaur from the Late Jurassic of North America supports a late acquisition of avian flight”. PeerJ, 7: e7247. https://www.ncbi.nlm.nih.gov/pmc/articles/PMC6626525/.
- David Lambert, The Dinosaur Data Book (New York: Avon Books, 1990), page 61; Don Lessem and Donald F. Glut, The Dinosaur Society Dinosaur Encyclopedia (New York: Random House, 1993), page 184; Peter Dodson, The Age of Dinosaurs (Lincolnwood: Publications International Ltd., 1993), page 142; Thomas R. Holtz Jr, Dinosaurs: The Most Complete, Up-To-Date Encyclopedia for Dinosaur Lovers of All Ages (New York: Random House, 2007), page 382.
- Thomas R. Holtz Jr, Dinosaurs: The Most Complete, Up-To-Date Encyclopedia for Dinosaur Lovers of All Ages (New York: Random House, 2007), page 147.
Bibliography
Books:
- Dodson, Peter. The Age of Dinosaurs. Lincolnwood: Publications International Ltd., 1993.
- Holtz Jr., Thomas R. Dinosaurs: The Most Complete, Up-To-Date Encyclopedia for Dinosaur Lovers of All Ages. New York: Random House, 2007.
- Lambert, David. The Dinosaur Data Book. New York: Avon Books, 1990.
- Lessem, Don; Glut, Donald F. The Dinosaur Society Dinosaur Encyclopedia. New York: Random House, 1993.
Articles:
- Fiorello Anthony R.; Adams Thomas L. (2012). “A therizinosaur track from the Lower Cantwell Formation (Upper Cretaceous) of Denali National Park, Alaska”. Palaios, 27: 395-400.
- Fiorillo, Anthony R.; McCarthy, Paul J.; Kobayashi, Yoshitsugu; Tomsich, Carla S.; Tykoski, Ronald S.; Lee, Yuong-Nam; Tanaka, Tomonori; Noto Christopher R. (August 3, 2018). “An unusual association of hadrosaur and therizinosaur tracks within Late Cretaceous rocks of Denali National Park, Alaska”. Scientific Reports, 2018; 8 (1) DOI: 10.1038/s41598-018-30110-8. https://www.nature.com/articles/s41598-018-30110-8.
- Harris, J. D.; Johnson, K. R.; Hicks, J.; Tauxe; L. (1996). “Four-toed theropod footprints and a paleomagnetic age from the Whetstone Falls Member of the Harebell Formation (Upper Cretaceous: Maastrichtian), northwestern Wyoming”. Cretaceous Research, 17: 381-401.
- Harris, J. D. (1997). “Four-toed theropod footprints and a paleomagnetic age from the Whetstone Falls Member of the Harebell Formation (Upper Cretaceous: Maastrichtian), northwestern Wyoming: a correction”. Cretaceous Research, 18: 139.
- Hartman, Scott; Mortimer, Mickey; Wahl, William R.; Lomax, Dean R.; Lippincott, Jessica; Lovelace, David M. (2019). “A new paravian dinosaur from the Late Jurassic of North America supports a late acquisition of avian flight”. PeerJ, 7: e7247. https://www.ncbi.nlm.nih.gov/pmc/articles/PMC6626525/.
- Lockley, Martin G.; Nadon, G.; Currie, Philip J. (2003). “A diverse dinosaur-bird footprint assemblage from the Lance Formation, Upper Cretaceous, eastern Wyoming; implications for ichnotaxonomy”. Ichnos, 11: 229-249.
- Lockley, Martin; Gierlinski, Gerard; Adach, Lidia; Schumacher, Bruce; Cart, Ken (2018). “Newly Discovered Tetrapod Ichnotaxa from the Upper Cretaceous Blackhawk Formation, Utah”. In Spencer G. Lucas and Robert M. Sullivan, eds. New Mexico Museum of Natural History and Science. Fossil Record 6, Volume 2: Bulletin 79. Albuquerque: New Mexico Museum of Natural History and Science, 2018. Pages 469-480.
- McCrea, R. T.; Buckley, L. G.; Plint, A. G.; Currie, Philip J.; Haggart, J. W.; Helm, C. W.; Pemberton, S. G. (2014). “A review of vertebrate track-bearing formations from the Mesozoic and earliest Cenozoic of western Canada with a description of a new theropod ichnospecies and reassignment of an avian ichnogenus”. In Lockley Martin G.; Lucas, Spencer G., eds. New Mexico Museum of Natural History & Science. Bulletin 62: Fossil Footprints of Western North America. Albuquerque: New Mexico Museum of Natural History & Science, 2014. Pages 5-94.
- McCrea, R. T.; Tanke, D. H.; Buckley, L. G.; Lockley, Martin G.; Farlow, James O.; Xing, L.; Matthews, N. A.; Helm, C. W.; Pemberton, S. G.; Breithaupt, B. H. (2015). “Vertebrate ichnopathology: pathologies inferred from dinosaur tracks and trackways from the Mesozoic”. Ichnos, 22 (3–4): 235-260.
- Zanno, Lindsay Elizabeth. A Taxonomic and Phylogenetic Reevaluation of Therizinosauria (Dinosauria: Theropoda): Implications for the Evolution of Maniraptora. PhD dissertation, submitted to the University of Utah. December 2008.
Websites:
- Alaska Dispatch. “Therizinosaur: prehistoric predator set standard for ‘weird’ in Alaska”, by Ned Rozell (September 29, 2012). http://www.alaskadispatch.com/article/therizinosaur-prehistoric-predator-set-standard-weird-alaska.
- Dinosaur Mailing List. “Re: Yet even more questions (and I’m sure there’ll be more…)”, by Mickey Mortimer (June 22, 2002). http://dml.cmnh.org/2002Jun/msg00369.html.
- Fossilworks. “Saurexallopus”. http://fossilworks.org/bridge.pl?a=taxonInfo&taxon_no=65843.
- Fossilworks. “Saurexallopus lovei”. http://fossilworks.org/bridge.pl?a=taxonInfo&taxon_no=65844.
- Fossilworks. “Saurexallopus zerbsti”. http://fossilworks.org/bridge.pl?a=taxonInfo&taxon_no=81011.
- National Park Service. “The Lower Cantwell Formation and Its Fossils”, by Alexander de Moor (2010). http://www.nature.nps.gov/geology/gip/web_products/DENA_2010_GIP_deMoor_Website.pdf.
- Science Daily. “First North American co-occurrence of Hadrosaur and Therizinosaur tracks found in Alaska” (August 6, 2018). https://www.sciencedaily.com/releases/2018/08/180806095216.htm?fbclid=IwAR0IpHB2CNwymxzEIw7bK8t4sOyalcbN3Tk_st2YBmaRSA6-5WI5gv9hqXk.
- Theropod Database. “Therizinosauroidea”. http://theropoddatabase.com/Therizinosauroidea.htm.
Videos:
- Utah’s Dino Graveyard. The Discovery Channel, 2005.
- When Dinosaurs Roamed America. The Discovery Channel, 2001.