Dinosaurs and Barbarians

Home » 2021 » February

Monthly Archives: February 2021

Ceratodus: The Iconic Lungfish of the Mesozoic Era

Ceratodus was a genus of prehistoric lungfish which existed on Earth for a surprisingly long time, from the late Triassic Period approximately 227 million years ago to the beginning of the Eocene Epoch of the Tertiary Period about 55 million years ago – a jaw-dropping span of 172 million years! That’s impressive by ANYBODY’S standards!

Lungfish as a whole are a primitive group of fish. They first appeared during the early Devonian Period about 416 million years ago (MYA), and it’s believed that they represent an evolutionary “missing link” between fish and amphibians. The closest relatives of the lungfish are the coelacanths, meaning “hollow spines”. That’s not surprising, considering that both lungfish and coelacanths have prehistoric origins as well as that both groups are classified as “lobe-finned fish”.

Lungfish do not have individual teeth like many fish today. Instead, they have four large bone plates (two in its upper jaw, and another two in its lower jaw) that were ridged in texture and crowned with thick triangular projections, and were used for crushing and cracking. Many species of modern lungfish feed on worms, freshwater snails, crustaceans, small fish, and amphibians.

Today, there are only six surviving species of lungfish, and all of them are found in hot tropical environments. With the exception of one species found in the Amazon Jungle and another species found in northern Australia, the remaining lungfish species are found in Africa.

  1. The South American Lungfish (Lepidosiren paradoxa), found in the Amazon River.
  2. The Marbled Lungfish (Protopterus aethiopicus), which is found throughout much of eastern and central Africa.
  3. The Gilled Lungfish (Protopterus amphibius), which is also found in eastern Africa.
  4. The West African Lungfish (Protopterus annectens), which is found, not surprisingly, in western Africa.
  5. The Spotted Lungfish (Protopterus dolloi), which inhabits the Congo Jungle of central Africa.
  6. The Australian Lungfish, also called the Queensland Lungfish (Neoceratodus forsteri), found in northeastern Australia. Of all of the extant lungfish species, this one is believed to be the most primitive.

Special attention must be given to the Australian Lungfish (Neoceratodus forsteri), for not only is this species regarded as the most archaic of all of the extant lungfish, but it was once believed to be the sole surviving member of the prehistoric lungfish genus Ceratodus alive in modern times.

Skeleton of Neoceratodus forsteri. From Günther, Albert. “Description of Ceratodus, a Genus of Ganoid Fishes, Recently Discovered in Rivers of Queensland, Australia”. Philosophical Transactions of the Royal Society of London, volume 161 (1871). Plate XXX. https://www.jstor.org/stable/pdf/109041.pdf.

The lower jaw of Neoceratodus forsteri, seen from above. From Krefft, Gerard. “Description of a gigantic amphibian allied to the genus Lepidosiren from the Wide-Bay district, Queensland”. Proceedings of the Zoological Society, volume 16 (April 28, 1870). Page 222. https://ia800405.us.archive.org/16/items/biostor-107043/biostor-107043.pdf.

The genus Ceratodus was established in 1837 by the famed Swiss ichthyologist Louis Agassiz based upon teeth which were found in European rock layers dated to the Triassic and Jurassic Periods. Most Ceratodus fossils that are found consist of isolated tooth plates, and different species have been named based largely upon difference in tooth morphology. Twenty-two species of Ceratodus have been named since the genus was first described in 1837. For a long time, Ceratodus was what is known as a “waste basket taxon” – all North American lungfish fossils were ascribed to this genus, regardless of how different they were from each other. Recently, a careful re-examination of lungfish fossils have revealed that these animals are remarkably different from each other and may constitute numerous genera, not just one. If that’s the case, then the overall lifespan of Ceratodus as a genus may be dramatically shorter than was previously supposed (Günther, Albert. “Description of Ceratodus, a Genus of Ganoid Fishes, Recently Discovered in Rivers of Queensland, Australia”. Philosophical Transactions of the Royal Society of London, volume 161 (1871). Page 512).

File:Ceratodus.jpg

Ceratodus, painted by Heinrich Harder. From Animals of the Prehistoric World (1916). Public domain image, Wikimedia Commons. https://commons.wikimedia.org/wiki/File:Ceratodus.jpg.

Ceratodus’ length varied depending on the species. Most sources which I have seen give an average length of 3 feet long. However, one species of Ceratodus may have reached truly gigantic proportions, possibly reaching 10 to 12 feet long. This estimate is based upon a single bone plate, which is the largest-known of any lungfish. The tooth plate was found in central Nebraska in rocks dated to the Miocene or Pliocene Epochs of the Tertiary Period. Shimada and Kirkland hypothesized that the tooth had been carried into central Nebraska by river from older rock layers that were located further to the west within Wyoming, in rocks dated to either the late Jurassic or early Cretaceous Periods. However, the tooth isn’t as banged up as you would expect from such a long journey. It’s possible that the tooth is endemic to central Nebraska, and if that is the case, 1) Ceratodus was alive in North America for a much longer geologic time span than previously supposed, or 2) This species is mis-identified and belongs to a new un-described genus of giant lungfish which lived in central North America about 5 million years ago, or 3) This was a species which happened to have unusually large teeth within its jaws, and the overall length of the animal was much smaller than the 4 meter estimate given by Shimada and Kirkland. Unfortunately, only one tooth plate has been discovered. Until more specimens are found, everything that we have to say about this specimen needs to be taken with a great degree of skepticism. (Kenshu Shimada and James I. Kirkland, “A Mysterious King-Sized Mesozoic Lungfish from North America”. Transactions of the Kansas Academy of Science, volume 114, issue 1 (2011). Pages 135-141. https://www.researchgate.net/publication/261964060_A_Mysterious_King-Sized_Mesozoic_Lungfish_from_North_America).

For the artwork accompanying this article, I decided to change up my style. For this drawing, I chose to evoke the whimsical style of the paleo-art of Patricia Bujard. If you don’t know who Patricia Bujard is, then I highly recommend that you check out her work. She is a children’s author and illustrator with a love for prehistoric life, and I find her artwork adorable. There aren’t too many people who can make an Allosaurus “cute”, but dag-nabbit, she somehow manages to pull it off. You can see her artwork on her WordPress page, Pete’s Paleo Petshop. My own drawing, which you can see below, was made with an ordinary Crayola black marker.

Ceratodus © Jason R. Abdale. February 9, 2021.

Keep your pencils sharp, and in this case, also keep your markers properly stored so they don’t dry out.

Promastodonsaurus

This is Promastodonsaurus, literally meaning “before Mastodonsaurus”. Despite its saurian name, it was not a dinosaur, or even a reptile. It was actually a large amphibian. Fossils of Promastodonsaurus were found in Argentina within the rocks of the Ischigualasto Formation, dated to the middle Triassic Period approximately 230 million years ago. The species was officially named in 1963 by the famed South American paleontologist José Bonaparté, in reference to another large amphibian named Mastodonsaurus which lived in Europe during a slightly later time.

Cladistically-speaking, this animal belonged to a large group of amphibians called the “labyrinthodonts”, so-named because a cross-section of their teeth looked like a maze. Within this broad group is a sub-division called the “temnospondyls”, “the cut vertebrae” because each of their backbones is divided into several parts. The temnospondyls were a diverse group of labyrinthodont amphibians which first appeared during the Carboniferous Period and lasted into the Cretaceous Period – a span of nearly 200 million years. Within the order Temnospondyli is the sub-order “Stereospondyli”, and within this is a division called the capitosaurians, “the head lizards”, so-named due to their freakishly huge heads. Promastodonsaurus was a member of this group. It was essentially a giant meat-eating salamander with the head of an alligator.

The only evidence that we have of this animal is a single partial skull. Based upon its similarity to the skulls of other temnospondyl amphibians within its family, it is believed that the animal’s head measured 45 centimeters long (Hans-Dieter Sues and Nicholas C. Fraser, Triassic Life on Land: The Great Transition. New York: Columbia University Press, 2010. Page 69). This in turn would make the entire animal somewhere in the vicinity of 6 feet long, as big as a medium-sized alligator.

Promastodonsaurus bellmani. © Jason R. Abdale. February 9, 2021.

During the middle Triassic Period, crocodilians did not exist, so the capitosaurians like Mastodonsaurus and Promastodonsaurus essentially filled in that ecological niche as crocodilian analogs. Large amphibians like these would continue to dominate freshwater environments until they were replaced by the phytosaurus, who in turn would be replaced by crocodilians.

 

History Lecture – “The Great Illyrian Revolt” at the Queens Public Library – January 26, 2021

Greetings everyone! On January 26, 2021, I conducted my first ever public lecture as a historian when I delivered a talk for the Queens Public Library via WEBEX concerning the Great Illyrian Revolt, a massive uprising which took place against the Roman Empire from 6 to 9 AD. The lecture was recorded on the host’s personal computer, and she sent me the link to the video, but I didn’t know how to download this video file onto my own computer until a few hours ago. After some very frantic computer work, here it is! The video lasts for a just a tad longer than an hour and twelve minutes. I hope you enjoy it.

If you like this lecture please purchase a copy of my book The Great Illyrian Revolt: Rome’s Forgotten War in the Balkans, AD 6-9, published by Pen & Sword Books in 2019.

February 2 – The Feast of Ceres and the Blessing of the Seeds

Today is February 2. Most Americans know this as “Groundhog Day” in which, according to their superstitions, a groundhog is able to predict if warm or cold weather will come. The story is that if a groundhog emerges from its burrow and sees its shadow, it will become frightened by it and scurry back inside. The act of returning to its burrow signifies that Winter is still here and it’s not practical to go outside looking for food just yet. However, if the groundhog exits its underground home and does not see its shadow, it will stay outside. This symbolizes that the groundhog is not afraid to venture out from its place of Winter hibernation and that Spring is right around the corner.

Many people nowadays scoff at such nonsense. How can an animal predict the coming of Spring? But did you know that there is an ancient religious background to this modern-day ritual? The beginning of February is important in a few religious calendars. It’s the time of the Feast of Ceres, the ancient Roman goddess of agriculture and the patron god of farmers; Ceres was the Roman version of the Greek goddess Demeter. It’s also the date for the ancient Celtic holy day of Imbolc. In ancient Ireland, the beginning of February was marked by a feast dedicated to the goddess Brigid, who was later turned into a Catholic Christian saint. Finally, early February marks the Catholic Christian festival of Candle Mass, which is either held in honor of the Virgin Mary or the day that the infant Jesus Christ was officially presented to the Jewish temple. This day was marked by a great lighting of candles in the churches and carried in processions, and it has been proposed that this ritual procession of people carrying candles originated in ancient Rome, because the Romans performed a nearly identical ritual in commemoration of the purification god Februus (1)

All three of these religious days have one thing in common: early February, either February 1 or 2, marks the half-way point between the coming of Winter and the coming of Spring. The beginning of February celebrates that Winter is half-way over, that the days are getting longer, and that warmer weather is on its way.

The ancient Roman seasonal calendar was different from ours. To us, the Vernal Equinox in late March marks the beginning of Spring, but to the ancient Romans, the Vernal Equinox marked the middle of Spring. To them, the real beginning of Spring was in early February, somewhere between February 1-4. Therefore, to the Romans, February 2 was their true Spring Festival.

Now that you know that warmer weather is coming, you need to get ready to plant your crops. In an earlier post, which you can read here, I described the ancient Roman agriculture festivals that took place in late January and early February. In that article, I stated that the Romans conducted a pair of religious rituals several days apart dedicated to the Mother Earth goddess Tellus and agriculture goddess Ceres. The Feast of Tellus, which was customarily observed on January 24 (although it didn’t always fall on this date), served to purify the soil by removing any pests and diseases in it and also called upon the Mother Earth goddess to bring good weather and other profitable growing conditions. The purification process would take several days, probably depending on how large your field was. Then on February 2, the second of the pair was celebrated, a festival dedicated to the goddess Ceres.

Like the first festival, the second was designed with the goal of purification. February 2 was marked by the Blessing of the Seeds, in which the Roman priests offered sacrifices of pork and wheat cakes, symbolizing livestock and crops, to Ceres and asked her to bless the seeds which they were about to plant into the ground and to remove any impurities from them, such as fungus, disease, or pests. Once all of the prayers and sacrifices had been made, the farmers planted the purified seeds into the purified soil (2).

The seeds which the Romans were about to plant had been kept in storage since October. It was prudent for farms to set aside the seeds that they intended to plant the following year. However, to keep these seeds safe, they needed to be locked away. If anything should happen to them, such as animals eating them or leaving them out to go moldy and rotten, there would be no food and the country would be gripped with famine. So, once the Autumn harvest was taken in, the Romans placed the following year’s seeds into underground storage containers, which were dedicated to Ceres. You can read more about this by clicking here (3).

The ancient Roman poet Ovid says that in the archaic past, the Romans would only offer grain and salt to their gods. However, they later added animal sacrifices to their rituals, and says that Ceres was the first god to have this done in her honor. On her feast day, a pig was sacrificed to her, supposedly as punishment for pigs uprooting a farmer’s crops. As Ovid relates, “Ceres was first to delight in the blood of the greedy sow, her crops avenged by the rightful death of the guilty creature. She learned that in Spring the grain, milky with sweet juice, had been uprooted by the snouts of bristling pigs. The swine were punished, terrified by that example” (4).

“Satisfy the eager farmers with full harvest, so they reap a worthy prize from their efforts. Grant the tender seeds perpetual fruitfulness, don’t let new shoots be scorched by cold snows. When we sow, let the sky be clear with calm breezes, sprinkle the buried seed with heavenly rain. Forbid the birds, that prey on cultivated land, to ruin the cornfields in destructive crowds. You too, spare the sown seed, you ants, so you’ll win a greater prize from the harvest. Meanwhile let no scaly mildew blight its growth, and let no bad weather blanch its colour, may it neither shrivel, nor be over-ripe and ruined by its own rich exuberance. May the fields be free of darnel that harms the eyesight, and no barren wild oats grow on cultivated soil. May the land yield rich interest, crops of wheat and barley, and spelt roasted twice in the flames” (5)

Note that Ovid makes NO MENTION of an agricultural festival occurring on February 2. All of the information that we have about this is mentioned in conjunction with the activities which took place on January 24.

Once all of the religious rituals had been conducted, the farmers could get to work sowing the seeds on their land. This was an important event in the minds of the ancient Romans. These were the first seeds planted in the year, and whether these seeds would do well or not would be a sign to the farmers about what the rest of the year would be like for them. If these first seeds did well, their Autumn harvest would be bountiful, but if they did poorly, they were likely going to suffer famine. In this way, we can see a similarity to the holiday of Groundhog Day – the good or bad fortune observed on this one day determines how things would play out in the future.

So on February 2, as you see snow blowing outside your window, be comforted in knowing that you are halfway towards sunshine and warmer days. Pray to Ceres for a good growing season for the coming Spring. Pray to Ceres to protect your seeds against birds, bugs, and bad weather. Offer sacrifices of pork and wheat cakes upon the household hearth.

Source citations

  1. John Audley, A Companion to the Almanack, 2nd Edition. London: 1804. Page 24; Reverend Alban Butler, The Lives of the Saints, Volume 2. Derby: Thomas Richardson & Son, 1798. Pages 34-42; Augustine Calmet, Calmet’s Great Dictionary of the Holy Bible, Volume II. Charlestown: Samuel Etheridge, Jr., 1813; Edward Augustus Kendal, Pocket Encyclopedia, or, A Dictionary of Arts, Sciences, and Polite Literature, Volume I. London: 1811. Page 329; Hipolito San Joseph Giral del Pino, A Dictionary: Spanish and English, and English and Spanish. London: 1763.
  2. Ovid, Fasti, book 1, January 24.
  3. William Warde Fowler, “Mundus Patet. 24th August, 5th October, 8th November”. Journal of Roman Studies, volume 2 (1912). Pages 25‑33; Thomas Morell and William Duncan, An Abridgement of Ainsworth’s Dictionary; English and Latin, Revised Edition. Edinburgh: Oliver and Boyd, 1862. Pages 29-30; Mark Bradley, “Crime and Punishment on the Capitoline Hill”. In Mark Bradley, ed., Rome, Pollution and Propriety: Dirt, Disease and Hygiene in the Eternal City from Antiquity to Modernity. Cambridge: Cambridge University Press, 2012. Page 120.
  4. Ovid, Fasti, book 1, January 9.
  5. Ovid, Fasti, book 1, January 24.

Bibliography

  • Audley, John. A Companion to the Almanack, 2nd Edition. London: 1804.
  • Bradley, Mark. “Crime and Punishment on the Capitoline Hill”. In Mark Bradley, ed., Rome, Pollution and Propriety: Dirt, Disease and Hygiene in the Eternal City from Antiquity to Modernity. Cambridge: Cambridge University Press, 2012.
  • Butler, Reverend Alban. The Lives of the Saints, Volume 2. Derby: Thomas Richardson & Son, 1798.
  • Calmet, Augustine. Calmet’s Great Dictionary of the Holy Bible, Volume II. Charlestown: Samuel Etheridge, Jr., 1813
  • Fowler, William Warde. “Mundus Patet. 24th August, 5th October, 8th November”. Journal of Roman Studies, volume 2 (1912). Pages 25‑33. http://penelope.uchicago.edu/Thayer/E/Journals/JRS/2/Mundus*.html.
  • Kendal, Edward Augustus. Pocket Encyclopedia, or, A Dictionary of Arts, Sciences, and Polite Literature, Volume I. London: 1811.
  • Morell, Thomas; Duncan, William. An Abridgement of Ainsworth’s Dictionary; English and Latin, Revised Edition. Edinburgh: Oliver and Boyd, 1862.
  • Pino, Hipolito San Joseph Giral del. A Dictionary: Spanish and English, and English and Spanish. London: 1763.
  • Ovid, Fasti, book 1, January 9. https://www.poetryintranslation.com/PITBR/Latin/OvidFastiBkOne.php.
  • Ovid, Fasti, book 1, January 24. https://www.poetryintranslation.com/PITBR/Latin/OvidFastiBkOne.php.

 

February 1 – The Month of Februus, the Ancient Roman God of Purification

In the ancient Roman calendar, several of the months are named after gods in the Roman pantheon. January is named after Janus, the god of new beginnings. March is named after Mars, the god of war. But what about February? February is the month of Februus, the god of purification. The name Februus comes from the Latin verb februa (which may have either Etruscan or Sabine roots), which means “to purge, purify, or cleanse”. The word “fever” is based on the same origin as the name “February”, because the Romans believed that you could purge sickness from your body by sweating it out of your system (1).

In addition to having the entire month dedicated in Februus’ honor, the ancient Romans also had a specific day dedicated to him in their calendar. This was called the Februalia, the Feast of Februus. In one source, it says that the Februalia purification ritual spanned from February 13 to 15 (2), but in all other sources that I have seen, it states that it only took place on the 15th. Later, the purification rituals of the Februalia were absorbed into the fertility ritual of the Lupercalia, which you can read about in more detail here.

In the past, the Roman calendar began with the month of March and ended with the month of February. Mars was seen as the divine father of the Romans, for he was the father of their first king Romulus; the year began with the month that bore his name. February, the final month of the calendar, was regarded as the death of the year, and consequently February was known for reverence to the dead (3). Traditionally, Roman government officials began their term-of-office on the first day of the year (March 1st) and exited on the last day of the year (February 28th or 29th). However, during the 150s to 130s BC, several important changes occurred within the Roman Republic, and many of these changes had to do with the Roman military campaigns in Spain. Military campaigns almost always began in March or April, when the temperature warmed, the snows melted, and the Roman Army could move. However, the immensities of the fighting in Spain meant that Roman military commanders were given precious little time to organize their campaigns. As a result of the wars in Spain, the rules and conventions of government needed to be changed due to the necessities of waging military operations in that theater, including extending the term-of-office for certain officials in order for them to more effectively carry out their duties, and even changing the Roman calendar. It was decided to shift the months of the calendar around. January and February, which had previously been the eleventh and twelfth months, were now moved to the beginning of the year to serve as the first and second months. Roman politicians and military commanders now assumed their powers on January 1 instead of March 1, which gave them at least sixty more days to prepare their troops for the upcoming military campaigns. This is also the reason why the months of September (literally translated to “Month Number 7”), October, November, and December now serve as the ninth, tenth, eleventh, and twelfth months of the year (4).

The Roman poet Ovid writes about the purification rituals of February in his Fasti:

“The fathers of Rome called purification ‘februa’. Many things still indicate that meaning for the word. The high priests ask the King and the Flamen for woolen cloths, called ‘februa’ in the ancient tongue. When houses are cleansed, the roasted grain and salt, the lictor receives, are called by the same name. The same name too is given to the branch, cut from a pure tree, whose leaves wreathe the priests’ holy brows. I’ve seen the priest’s wife (the Flaminica) ask for ‘februa’, and at her request she was given a branch of pine. In short anything used to purify our bodies, had that title in the days of our hairy ancestors. The month is so called, because the Luperci cleanse the earth with strips of purifying hide, or because the time is pure, having placated the dead, when the days devoted to the departed are over. Our ancestors believed every sin and cause of evil could be erased by rites of purification. Greece set the example: she considered the guilty could rid themselves of sins by being purified” (5)

Ovid also states that a gathering was held every February 1 at the Forest of Alernus, located where the Tiber River empties into the Mediterranean Sea. Unfortunately, he does not provide any reason for this assembly of people at this wood, nor does he go into any description as to what activities occurred there. Elsewhere, at the tomb of Numa Pompilius and at Jupiter’s temple of the Capitoline Hill, a sheep was sacrificed. (6)

Ovid also states that February 1 marked the date that at least two temples were built dedicated to the goddess Juno, the queen of the Roman gods. However, Ovid remarks that these temples had long fallen into ruins by the time that he was writing (7).

Source citations

  1. Definitions and Translations. “Februa”. https://www.definitions.net/definition/februa.
  2. Definitions and Translations. “Februa”. https://www.definitions.net/definition/februa.
  3. Ovid, Fasti, book 2, introduction.
  4. Rome and the Barbarians, lecture 8 – “The Roman Conquest of Spain”.
  5. Ovid, Fasti, book 2, introduction.
  6. Ovid, Fasti, book 2, February 1.
  7. Ovid, Fasti, book 2, February 1.

Bibliography